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Index >>Bacterial Fungal Cells>>Flagella and Fimbria

Flagella and Fimbria

Flagella and Fimbria  -

The presence of flagella on some bacteria has been known since the early part of the 19th century and both motility and the form of flagellation have been used as a taxonomic tool in bacterial classification.

Flagella are surface appendages found in motile bacteria and appear as filaments usually of 12-20 ,nm in diameter, and about 6.8 micrometer in length.

The diameter of the individual flagellum in a culture is usually constant but the length may vary.

The location of the flagella in different bacteria varies and maybe, either polar, sub polar, or peritrichous the, number of flagella per, cell also varies with the bacterial species.

The flagellar apparatus is composed of three distinct parts namely the filament, the hook and the basal granule.

The outermost structure is the filament which is a fibre composed of a protein known as flagellin (subunit molecular weight 20,000) and this is attached to the basal granule with the aid of the hook.

The hook and the basal granules contain proteins which are antigenically distinct from the flagellin.

The basal body consists of a small central rod inserted in to a system of rings and the unit operates basically as a simple motor.

Growth of the flagella occurs by the addition of the flagellin subunits at the distal end after being carried through from the cytoplasm, apparently through the hollow core of the flagellum.

Flagella are involved in bacterial motility. Mechanical deflagellation makes motile cells immotile.

Movement of the bacterial cell occurs by the rotation of the flagella either clockwise or anti clockwise along its long axis.

The cell has the capacity to vary both the speed and the direction of rotation as well as the frequency of stops and starts by appropriate movement of the flagella.

Peritrichously flagellated bacteria swim in a straight line over moderate distance, These runs are interrupted periodically by sudden changes in direction leading to tumbling.

The movement of the bacteria, is therefore said to be zig zag.

It is found that smooth swimming in a fixed direction is mediated b)' the rotation of flagella in an anticlockwise direction and tumbling is caused by the rotation in a clockwise direction.

In bacteria with polar flagella, a change in the direction occurs by the change in the direction of rotation.

The pili or the fimbriae are filamentous hair like structures on the surface of only Gram negative bacteria and are distinct from the flagella.
The term fimbriae is used for all hair like structures other than the structures involved in transfer of genetic material which are termed as pili.

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