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Index >> Bacterial Photosynthesis >> Chromatiaceae (Formerly Thiorhodaceae ) Purple Sulphur Bacteria

Chromatiaceae (Formerly Thiorhodaceae ) Purple Sulphur Bacteria

Chromatiaceae (Formerly Thiorhodaceae ) Purple Sulphur Bacteria - The purple sulphur bacteria form globules of elemental sulphur from sulphide inside cells. There is one photocentre. Hydrogen sulphide, sulphur, thiosulpbate or molecular hydrogen are sources of hydrogens.

Organic molecules are utilized in some cases. The principal photosynthesis pigments are bacteriochlorophylls a and b which show absorbtion bands at 375 and 590 nm in the visible region and 800, 850 and 890 in the infrared region for their chlorophylls. A large amount of carotenoids of groups 1, 3 and 4 or tetrahydrospirilloxanthin are present.

The absorption peaks of the carotenoids are between 400 and 600 nm.The primary photochemical reaction centre from the bacteriochlorophyll in Chromatium chromatophores is P890 (equivalent to P870 in photosynthetic strains). P800, is the light-harvesting cytochrome, c553, related to or identical with c555 of Chlorobium.

An additional cytochrome, cc', complexes, with P870. Coenzyme Q7 is associated with the reaction centre. In addition to the cyclic flow of electrons described above, a non cyclic electron transport system has also been suggested, but the evidence is conflicting. The noncyclic system, including a P905 reaction centre and cytochrome c552, would function at high light intensities or under more reduced conditions.

It can only operate under anaerobic conditions. The P890 pathway requires more oxidizing conditions and low light intensities. The existence of a second distinct photochemical centre (P905) bas been questioned.

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