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Index >> Soil, Nature Medium For Plant Growth >> Soil Structure

Soil Structure

Soil Structure

Movement of air and water in soil is governed by the structure of soil. The structure of soil, in turn, is dependent on stable aggregates of soil particles. A soil aggregate is a naturaIIy occurring cluster of soil particles with a strong binding force among them. Soil organic matter including humus, polysaccharides and polyuronides produced by soil microorganisms help to cement soil particles together while filamentous fungi provide additional mechanical support.

Several workers have found that the aggregating influence of different groups of soil microorganisms can be graded in the order, fungi > strep tomyces > gum producing bacteria > yeasts. Rapidly growing Rhizopus, Mucor, Chaetomium, Fusarium, Cladosporium, Aspergillus and Rhizoctonia are good examples of fungi known to secrete gums and provide mechanical support to bind soil particles. Examples of bacteria producing significant amounts of gum are Azotobacter, Beijerinckia, Rhizobium, Xanthomonas and Bacillus.

The lipopolysaccharides from gums of Rhizobium and Agrobacterium have been analysed and found to contain glucose, rhamnose, mannose, glucosamine and 4-0-methylglucoronic acid as major constituents. Other components of the gums are fructose, arabinose, galactose and xylose. It has been claimed that infection of root hairs of legumes is dependent on a water soluble polysaccharide of the rhizobial gum, although no conclusive evidence has been provided to support such a hypothesis.

Different soil components have been chemically extracted from soil and their effect on soil and their effects on soil aggregation properties studied. Among the carbohydrates isolated from soil, dextran containing high amounts of uronic acid which is resistant to microbial degradation has been shown to have the best soil aggregating qualities.

Several experiments have been carried out under laboratory conditions to demonstrate the effect of single or mixed cultures of microorganisms in soil amended with various carbon sources. The results of such experiments, however fascinating they may be, must be viewed with caution. It is also interesting to note that the involvement of polysaccharides in soil aggregation has been questioned by some workers based on results of experiments in which native polysac­charides of soil samples were destroyed by chemical treatments without any detriment to the property of soil aggregation.

 

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